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The Agronomic and Commercial Dynamics of Starke Ayres Spinach Germplasm in the East African Horticultural Sector

Kilimokwanza.org Report

1. Executive Landscape and Historical Genesis

The narrative of Kenya’s horticultural development is intrinsically linked to the evolution of its seed systems. Within this complex ecosystem, the cultivation of leafy green vegetables serves as a critical pillar of both national food security and the economic sustenance of smallholder farmers. Among these crops, the vegetable colloquially known as “spinach” occupies a dominant position in peri-urban and rural agriculture. However, the market is characterized by a distinct botanical dichotomy: the widespread cultivation of Swiss Chard (Beta vulgaris var. cicla) for the mass domestic market, and the niche, high-value production of True Spinach (Spinacia oleracea) for export and premium retail channels.

Starke Ayres, a seed enterprise with deep historical roots in Southern Africa, has established itself as a pivotal player in this landscape. To understand the company’s current market dominance and strategic trajectory in Kenya, one must first examine its corporate lineage. Tracing its origins back to 1877 in Cape Town, South Africa, the company was founded by Charles Ayres as a nursery and florist, later merging with Charles Starke’s grain and produce business.1 This union created a legacy that spans over 140 years, evolving from a localized seed merchant into a continental powerhouse under the aegis of the Plennegy Group.2 The Plennegy Group’s ownership structure is significant; as a family-owned conglomerate with over 3,500 employees serving the entire agricultural value chain, it provides Starke Ayres with the vertical integration and financial stability necessary to engage in long-term genetic research rather than chasing quarterly profits.2

The company’s expansion into the African continent was a calculated strategic maneuver designed to mitigate the risks associated with a single-market focus. While the South African home base remains critical, the operational footprint began to widen significantly in the late 20th century. A subsidiary was opened in Namibia in 1998, marking the first concrete step of expansion outside South Africa.2 This was subsequently followed by the establishment of subsidiaries in Zambia and Kenya. The entry into Kenya was particularly pivotal. Initially serving the market through distributors in the early 1990s, the demand for high-quality genetics necessitated the establishment of Starke Ayres Kenya Limited, a fully-fledged subsidiary.3

The Kenyan subsidiary is strategically located in Embakasi, Nairobi, along the critical Mombasa Road corridor.4 This location is not arbitrary; it situates the company at the logistical heart of East Africa, adjacent to the Jomo Kenyatta International Airport for airfreight exports and on the main arterial route connecting the port of Mombasa to the agricultural hinterlands of the Rift Valley and Western Kenya. From this hub, Starke Ayres manages a distribution network that not only permeates the Kenyan counties but also influences the broader East African Community, including Uganda, Tanzania, and Rwanda.5

A defining characteristic of Starke Ayres’ history in Kenya is its strategic pivot regarding its target demographic. Historically, the Kenyan vegetable seed market was bifurcated: high-cost, high-performance hybrids were imported almost exclusively for export growers producing French beans, snow peas, and baby spinach for European supermarkets, while the domestic market relied on low-cost, Open Pollinated Varieties (OPVs) often saved from previous harvests. In the past, 70% of Starke Ayres’ Kenyan turnover was derived from these large-scale export growers.7 However, recent years have witnessed a fundamental inversion of this ratio. Current data indicates that smallholder farmers now contribute approximately 55% of the company’s turnover, with export growers accounting for 40%.7

This shift is driven by powerful demographic and economic forces. The World Economic Forum predicts that Africa’s urban population will triple to 1.34 billion by 2050, creating an unprecedented demand for food within the continent’s own borders.5 Starke Ayres recognized that the smallholder farmer is the engine of this domestic food security. Consequently, the company reoriented its breeding and marketing strategies to empower these growers with “premium” genetics—varieties that are not just high-yielding but are robust enough to withstand the logistical realities of the developing world. This includes breeding for traits such as thicker leaf cuticles to survive transport on bumpy rural roads, a critical factor for perishable greens like spinach and chard.7

2. Global Genetic Integration and Breeding capabilities

The competitive advantage of Starke Ayres in the spinach sector is not solely derived from its local presence but from its integration into the global network of plant breeding. The modern seed industry is characterized by consolidation and technological partnership, and Starke Ayres has navigated this landscape to secure access to world-class germplasm.

2.1 The Plennegy-Pop Vriend-KWS Nexus

A critical element of the Starke Ayres spinach portfolio is its relationship with global breeding giants. While Starke Ayres conducts extensive proprietary breeding, the global standard for spinach genetics has long been set by Dutch breeders. Specifically, Pop Vriend Seeds, a family-owned Dutch company headquartered in Andijk, historically dominated the global market for spinach, beans, and Swiss chard seeds.8

In June 2019, a seismic shift occurred in the vegetable seed industry when KWS SAAT SE, a German plant breeding giant, acquired Pop Vriend Seeds for a price equivalent to approximately 20% of KWS’s market capitalization.8 This acquisition was significant for the African market because Starke Ayres and Pop Vriend have historically shared a synergistic relationship. The acquisition by KWS integrated Pop Vriend’s massive library of spinach germplasm—specifically their advanced resistance packages against Downy Mildew—into a larger, more resource-rich corporate structure.8

For the Kenyan farmer, this high-level corporate maneuvering translates directly into seed quality. Starke Ayres leverages these global connections to introduce varieties like Dash F1, which contain resistance genes developed in the high-pressure disease environments of Europe and North America. The partnership allows Starke Ayres to offer varieties that are tested in diverse ecological zones; while applied breeding is concentrated at six research stations in South Africa, the company maintains breeder trial sites in the USA, Argentina, Brazil, Hungary, and New Zealand.10 This global testing grid ensuring that a variety released in Kenya has likely already proven its resilience against a wide array of abiotic and biotic stressors.

2.2 Technological Sophistication in Breeding

The transition from phenotypic selection (selecting the best-looking plant) to genotypic selection (selecting the best DNA) marks the modern era of Starke Ayres’ R&D. The company employs advanced biotechnological tools including Marker-Assisted Breeding (MAB) and Double Haploid technology.10

  • Marker-Assisted Breeding (MAB): This technology allows breeders to identify specific genes linked to desirable traits—such as resistance to a specific race of Downy Mildew or tolerance to bolting—without waiting for the plant to fully mature and exhibit the trait. This significantly accelerates the breeding cycle, allowing Starke Ayres to respond rapidly to new pathogen strains emerging in East Africa.
  • Double Haploid Breeding: This technique is used to produce completely homozygous (genetically uniform) lines in a single generation, bypassing the years of inbreeding required in traditional methods. This ensures that when a Kenyan farmer buys a packet of hybrid spinach seed, every single seed is genetically identical, leading to the extreme crop uniformity required by supermarkets.10

Glendon Ascough, the Research Director at Starke Ayres, has emphasized the evolution of the company from a historic seed merchant to Africa’s largest independent vegetable breeding company.11 His focus on “data-driven” breeding involves collaboration with entities like Computomics to use machine learning for predicting hybrid performance. While initially applied to crops like sweet corn, the mindset of using high-quality, diverse phenotypic data to predict performance is applicable across the portfolio, including leafy greens.11 This approach minimizes the risk of releasing a variety in Kenya that might fail under specific local conditions, such as the distinct photoperiods found at the equator.

3. Botanical Taxonomy and Nutritional Positioning

To conduct a precise analysis of the Kenyan market, it is imperative to navigate the botanical confusion that persists in the region. The term “Spinach” is used as a catch-all commercial term, but it encompasses three distinct botanical entities within the Starke Ayres catalogue.

3.1 Beta vulgaris var. cicla (Swiss Chard)

This is the “Spinach” of the masses in Kenya. It is technically a beet grown for its leaves rather than its root. The Starke Ayres catalogue explicitly clarifies this relationship, noting that Swiss Chard belongs to the beet family and has been cultivated in Europe for thousands of years.12 In South Africa and Kenya, it acts as a spinach substitute because it is far more heat-tolerant and higher-yielding than true spinach.

The dominance of Swiss Chard is driven by its physiology. It possesses large, fleshy leaves with a robust midrib. Unlike true spinach, which bolts (flowers) rapidly under the heat and long days of the tropics, Swiss Chard is a biennial that can be harvested continuously for months. Starke Ayres positions varieties like Fordhook Giant and STAR 1801 in this category.13

3.2 Spinacia oleracea (True Spinach)

This is the “Baby Spinach” or “English Spinach” found in premium salad mixes. It is a distinct species that is far more sensitive to environmental stress. It requires cooler conditions and is highly susceptible to soil-borne diseases and foliar pathogens. Starke Ayres services this market with varieties like Dash F1.15 The distinction is critical because the agronomic management for Spinacia oleracea—requiring high density sowing and strict irrigation control—is fundamentally different from the robust, spaced planting of Swiss Chard.

3.3 Amaranthus spp. (Amaranth/Marog)

Starke Ayres also lists Amaranthus (often called “Marog” or “Terere” in Kenya) within its leafy green portfolio.1 While often considered a traditional or indigenous vegetable, Starke Ayres has commercialized it, recognizing its status as a “superfood.” The catalogue highlights its high concentration of antioxidants and phytosterols, positioning it not just as a subsistence crop but as a nutraceutical product.1 The inclusion of Amaranth demonstrates Starke Ayres’ responsiveness to the “Indigenous Vegetable” renaissance in Kenya, where consumers are increasingly seeking traditional crops for their perceived health benefits.

3.4 Nutritional Marketing Strategy

Starke Ayres explicitly utilizes nutritional data as a marketing tool. Their catalogues describe Swiss Chard and Amaranth as “superfoods” loaded with Vitamin K, A, C, magnesium, and manganese.1 This alignment with the “wellness” trend is strategic. By framing these crops as essential for “skin, hair, and bone health” and capable of “lowering blood pressure” 1, they appeal to the growing urban middle class in Nairobi who are becoming as health-conscious as their counterparts in Europe. This marketing narrative helps justify the premium price of their seeds compared to generic alternatives.

4. Detailed Variety Performance and Agronomic Analysis

The core of Starke Ayres’ value proposition lies in the specific performance characteristics of its flagship varieties. A comparative analysis reveals how these varieties address the specific pain points of Kenyan growers.

4.1 Fordhook Giant: The Market Sovereign

Fordhook Giant is the ubiquitous Swiss Chard variety in East Africa. While it is an open-pollinated variety (OPV) widely available from various breeders (including Simlaw Seeds and Kenya Seed Company), Starke Ayres markets a specific strain that is selected for superior vigor and uniformity.

  • Morphology: The variety is characterized by large, dark green, crinkled (savoy) leaves with broad, glossy white midribs.13 The “savoy” texture is not merely aesthetic; the crinkles increase the surface area of the leaf, potentially enhancing photosynthetic capacity per unit of ground area. The broad white stem is a key quality indicator for the fresh market, where consumers associate the crisp whiteness with freshness.
  • Yield Metrics: Agronomic data suggests a yield potential of 10–12 tons per acre under standard smallholder conditions.16 However, intensive management can push this significantly higher. Competitor catalogues (Simlaw) list potential yields for Fordhook Giant at up to 20 tons per acre.17 The discrepancy likely stems from Starke Ayres’ conservative estimates based on marketable “bunched” yield versus total biomass.
  • Maturation and Cycle: The variety matures in 40–60 days after sowing.18 This rapid turnover is crucial for cash flow. A farmer can technically fit 5-6 distinct crop cycles in a year, although in practice, the crop is often managed as a “cut-and-come-again” perennial, where the outer leaves are harvested continuously while the central growing point remains intact.
  • Market Position: Starke Ayres emphasizes the “dark green” color of their strain. In the Kenyan market, darker leaves are often perceived by consumers as being more nutritious and less fibrous than paler variants.

4.2 STAR 1801: The Commercial Hybrid Challenger

As the market matures, Starke Ayres is pushing the adoption of STAR 1801, a hybrid Swiss Chard that represents a technological leap over Fordhook Giant.

  • Frilly Leaf Architecture: STAR 1801 features a “frilly” or deeply curled leaf margin.14 This morphology has a distinct commercial advantage: it creates volume in the bunch without adding proportionate weight. When sold by the bunch (volume) rather than by the kilogram, this allows farmers and retailers to create visually impressive displays. Furthermore, the frilly texture prevents the leaves from matting together when packed in sacks for transport, improving airflow and reducing the “heat of respiration” that causes rotting during transit to markets like Wakulima or Kongowea.
  • Shelf-Life Extension: The variety is bred with thicker leaves, directly addressing the post-harvest losses that plague Kenyan horticulture. The increased shelf-life 19 allows the produce to remain turgid on the supermarket shelf for longer periods, reducing waste for retailers like Naivas and Carrefour.
  • Processing Efficiency: STAR 1801 is noted for having a “low fibre white midrib”.14 Low fiber content in the stem makes the vegetable easier to cut and more palatable, addressing a common consumer complaint about older Swiss Chard stems being stringy.
  • Yield Stability: As a hybrid, STAR 1801 offers extreme uniformity. Every plant in the field grows at the same rate and reaches the same size. This uniformity facilitates synchronized harvesting, which is essential for commercial growers supplying large contracts who need to predict their tonnage accurately.

4.3 Dash F1: The Export-Grade Specialist

Dash F1 is a true baby spinach (Spinacia oleracea) hybrid designed for the most sophisticated segment of the market.

  • Speed of Production: This variety is an agronomic sprinter, maturing in just 30–35 days.21 This allows for incredibly rapid crop cycles, potentially enabling 8-10 rotations per year in a controlled environment.
  • Growth Habit: Dash F1 exhibits an upright growth habit.21 In mechanical or hand harvesting, this is a critical trait. Upright leaves stay off the soil, significantly reducing the accumulation of grit and soil splashes on the foliage. This reduces the washing costs for processors and lowers the incidence of soil-borne bacterial infections entering the leaf canopy.
  • Genetic Resistance (The RPF3 Locus): The most vital characteristic of Dash F1 is its disease resistance. True spinach is plagued by Downy Mildew (Peronospora effusa). Starke Ayres, leveraging global breeding advancements, incorporates resistance genes. Advanced genomic research has mapped resistance loci, such as the RPF3 locus, to specific chromosomes (Chromosome 3) near NBS-LRR disease resistance genes.22 While the specific race resistance of Dash F1 is proprietary, the “High Resistance” (HR) designation usually implies coverage against modern races (Pe: 1-15+). Without this genetic firewall, growing baby spinach in Kenya’s humid highlands would be an exercise in futility.

5. Environmental Interactions and Agronomic Protocols

The performance of Starke Ayres seeds is inextricably linked to the environmental conditions of Kenya. The company provides detailed protocols to bridge the gap between genetic potential and realized yield.

5.1 Soil Chemistry and Nutrition

Swiss Chard is notoriously sensitive to soil acidity. Starke Ayres production guidelines explicitly warn farmers to avoid soils with a pH below 6.0.12 In many parts of Kenya, particularly the tea-growing zones of the central highlands and parts of Western Kenya, soils are naturally acidic (pH 4.5 – 5.5). At this acidity, aluminum toxicity stunts root growth, and essential nutrients like magnesium become chemically locked out.

  • Remediation: The company advises applying agricultural lime 2-3 months before planting to rectify this acidity.12 This lead time is crucial because lime reacts slowly in the soil; applying it at planting is ineffective for that season’s crop.
  • Fertilization: Swiss Chard is a heavy feeder, particularly of Nitrogen (for leaf growth) and Potassium (for leaf turgidity and disease resistance).
  • Alternative Nutrition: Recent studies in Kenya have evaluated the use of Black Soldier Fly Frass Fertilizer (BSFFF) on Swiss Chard. Research indicates that integrating BSFFF with mineral fertilizers (NPK) under daily irrigation regimes significantly enhances yield and chlorophyll concentration.23 This is highly relevant for urban farmers in Nairobi using Starke Ayres seeds in sack gardens, where organic nutrient sources like insect frass are increasingly available and affordable compared to synthetic inputs.

5.2 Climatic Suitability and Stress Tolerance

  • Temperature Thresholds: The optimal temperature range for Starke Ayres Swiss Chard is 12–24°C.12 This perfectly matches the climate of Kenya’s high-altitude zones (1500m – 2500m ASL).
  • Heat Stress: While the crop prefers cool weather, varieties like Fordhook Giant possess a degree of heat tolerance. However, the company notes that temperatures above 32°C can degrade leaf quality.12 Under such heat, the plant prioritizes survival over vegetative growth, often leading to bitter leaves or bolting.
  • Frost Tolerance: The varieties can tolerate light frost, which is common in Limuru and Nyahururu during July and August.12 However, persistent heavy frost will damage the cellular structure of the leaves, rendering them unmarketable.

5.3 Plant Population and Spacing

Optimizing plant density is key to maximizing revenue. Starke Ayres recommends a plant population of 250,000 to 280,000 plants per hectare for direct drilling.12

  • Spacing: The recommended spacing is 10-15 cm within the row and 450-500 mm between rows.12
  • Canopy Management: Correct spacing allows for rapid canopy closure. As noted by Kenyan farmers, a closed canopy shades out weeds, reducing labor costs for manual weeding and maintaining soil moisture.25 This concept of “living mulch” is a critical component of the agronomic advice embedded in Starke Ayres’ technical support.

6. Commercial Value Chain and Economic Implications

6.1 The Economics of Premium Seed

The shift from OPVs to hybrids (like STAR 1801) or premium OPVs (Starke Ayres Fordhook Giant) entails a higher upfront cost for the farmer. Starke Ayres justifies this premium through germination reliability and risk mitigation.

  • The Cost of Failure: For a smallholder, a packet of seeds that fails to germinate is a catastrophic loss of time, not just money. Starke Ayres operates ISTA (International Seed Testing Association) accredited laboratories.3 This accreditation guarantees that the germination rates stated on the packet are scientifically verified. A 95% germination rate versus a competitor’s 75% drastically alters the “cost per established plant.”
  • Uniformity as a Value Driver: Uniformity is the currency of the modern supermarket supply chain. Retailers demand bunches that look identical. Hybrid seeds produce phenotypes that are genetically identical, allowing farmers to harvest an entire bed at once with consistent sizing. This reduces the labor cost of sorting and grading, which often eats into profit margins.

6.2 Bulk Seeds for Commercial Scale

Recognizing the duality of the market, Starke Ayres has introduced “bulk seed” options for commercial farmers.26 Selling seeds in larger, bulk packaging reduces the unit cost per seed compared to the small foil sachets sold to home gardeners. This strategy aligns with the needs of medium-scale farmers in counties like Kirinyaga and Kajiado who are planting acres rather than garden beds. It also supports sustainable practices by reducing the plastic waste associated with small-format packaging.26

6.3 Distribution and Logistics

The distribution network of Starke Ayres is a strategic asset. By partnering with major regional distributors like Holland Greentech, Twiga Chemicals, and Elgon Kenya 27, the company ensures that its seeds are available deep in the rural interior.

  • Supply Chain Resilience: During the COVID-19 pandemic, Starke Ayres was designated as an “Essential Service” provider.2 This classification was crucial. It allowed the company to continue moving seed stock across borders and county lines during lockdowns, proving the resilience of their supply chain. This reliability cemented the loyalty of many farmers who saw other supply chains collapse during the crisis.
  • Counterfeit Mitigation: The robust formal distribution network also acts as a buffer against counterfeit seeds—a rampant problem in Kenya. By selling through authorized stockists and “Agrovets,” Starke Ayres can better guarantee the integrity of the genetic material reaching the farmer.

7. Pathology and Biotic Risk Management

Disease pressure is the primary yield-limiting factor in Kenyan horticulture. The “Spinach Complex” is under constant siege from fungi and bacteria, particularly during the “Long Rains” (March-May) and “Short Rains” (October-December).

7.1 The Fungal Threat

Kenyan farmers report high susceptibility of Swiss Chard to fungal diseases during wet seasons.25 Common pathogens include Cercospora leaf spot, which causes unsightly blemishes that render the leaves unsellable.

  • Morphological Defense: The upright growth habit of STAR 1801 and Dash F1 is a morphological defense mechanism. By keeping the leaves more vertical, airflow is improved within the canopy, allowing leaves to dry faster after rain or irrigation.21 Dry leaves are far less hospitable to fungal spores than wet ones.

7.2 The Downy Mildew Arms Race

For True Spinach (Dash F1), the battle is against Peronospora effusa. This pathogen evolves rapidly, with new races appearing regularly. Recent years have seen the denomination of new races Pe: 18 and Pe: 19, which have broken the resistance of older varieties.29

  • The Hybrid Solution: Starke Ayres’ access to the KWS/Pop Vriend germplasm library is critical here. Pop Vriend has released varieties with resistance up to Pe: 1-20.30 While Starke Ayres simplifies its marketing to “Disease Resistance,” the underlying genetics likely encompass these advanced race-specific resistance genes. This protects the farmer’s investment against a pathogen that can wipe out a crop in 48 hours.

8. Future Outlook and Emerging Trends

8.1 Vertical and Sack Farming

Urbanization is driving a revolution in “Sack Farming” and vertical gardens in Nairobi’s informal settlements and apartment balconies. Research utilizing the “Wonder Multistorey Gardening System” has shown that Swiss Chard is exceptionally well-suited to these intensive systems.23 Starke Ayres is well-positioned to dominate this niche. Their varieties, particularly the compact and vigorous hybrids, respond well to the high-input, high-output nature of vertical farming. The integration of organic nutrient solutions (like insect frass) with Starke Ayres genetics offers a sustainable model for urban food security.

8.2 Climate Resilience and Bolting Tolerance

Climate change in East Africa is manifesting as erratic rainfall and higher average temperatures. 85% of farmers already perceive these temperature shifts.31 Heat triggers bolting (premature flowering) in spinach and chard. Future breeding efforts at Starke Ayres are heavily focused on “slow bolting” traits.32 By developing varieties that remain in the vegetative phase even under heat stress, Starke Ayres effectively extends the growing season for farmers in semi-arid areas like Machakos and Kitui.

8.3 Data-Driven Breeding

The future of Starke Ayres in Kenya is digital. The collaboration with Computomics to use machine learning for phenotypic prediction 11 signals a move away from trial-and-error breeding. By modeling how specific genetic combinations will perform under the specific rainfall and temperature patterns of East Africa, Starke Ayres can design “climate-smart” varieties in silico before testing them in the field. This reduces the time-to-market for new varieties, ensuring that Kenyan farmers have access to genetics that are adapted to the climate of today, not the climate of ten years ago.

Conclusion

Starke Ayres has effectively cemented its status in the Kenyan horticultural sector by bridging the gap between advanced global breeding technologies and the practical realities of African smallholder farming. The company has successfully transitioned from an export-oriented supplier to a partner in domestic food security, recognizing the immense potential of the smallholder sector.

The portfolio strategy is clear and effective: Fordhook Giant remains the workhorse, providing reliable, high-volume nutrition for the mass market. STAR 1801 introduces the benefits of hybridization—uniformity, shelf-life, and disease tolerance—to the mainstream chard market, facilitating the professionalization of farmers targeting supermarkets. Dash F1 serves the elite tier, bringing world-class disease resistance to the high-risk, high-reward baby spinach sector.

For the agricultural stakeholder in Kenya—whether a farmer, investor, or policymaker—Starke Ayres represents a benchmark of quality. Their seeds function as a technology transfer mechanism, packaging complex genomic research into a product that requires no special equipment to use, only the soil and labor of the farmer. As the pressures of urbanization, climate change, and supermarket consolidation intensify, the “clean,” “uniform,” and “resilient” genetics provided by Starke Ayres are positioned to become not just a premium option, but the industry standard.


Appendix: Technical Product Reference & Agronomic Guidelines

Product NameSpeciesCategoryKey FeatureRecommended DensityResistance / ToleranceTarget Market
Fordhook GiantBeta vulgarisOPV Swiss ChardLarge white rib, heavy yield, robust250k – 280k / haLight Frost, HeatWet Markets, Schools, Institutions
STAR 1801Beta vulgarisHybrid Swiss ChardFrilly leaf, long shelf life, low fibre250k – 280k / haBolting tolerance, Fungal toleranceSupermarkets, Greengrocers
Dash F1Spinacia oleraceaHybrid Baby SpinachFast maturity (30-35 days), uprightHigh Density (Drill)Downy Mildew (P. effusa)Export, High-end Hotels, Salad Mixes
Green WaveBeta vulgarisOPV Swiss ChardAlternative textureStandardGeneral VigorHome Gardens, Local Market

Agronomic Protocol Summary

  • Sowing Depth: 2 cm.
  • Spacing: 10-15 cm in-row; 450-500 mm between rows.
  • Soil pH: > 6.0 (Lime required if acidic).
  • Temperature: Optimal 12-24°C.
  • Irrigation: Critical during germination; avoid waterlogging to prevent soft rot.
  • Harvest: “Cut-and-come-again” for Swiss Chard; single cut for Baby Spinach.

Citations:

Official Company Websites & Publications

Affiliated or Partner Organizations

Academic & Research Sources

  • Kabaru, J. et al. (n.d.). Agronomic Performance of Kale (Brassica oleracea) and Swiss Chard (Beta vulgaris) Grown on Soil Amended with Black Soldier Fly Frass Fertilizer under Wonder Multistorey Gardening System. The Hive, icipe. https://thehive.icipe.org
  • Kabaru, J. et al. (n.d.). Agronomic Performance of Kale and Swiss Chard… ResearchGate. https://www.researchgate.net
  • Quesada-Ocampo, L. M., et al. (n.d.). Denomination of Pe: 18 and 19, two new races of downy mildew in spinach. University of California Agriculture and Natural Resources (UC ANR). https://ucanr.edu
  • Rashed, A. et al. (n.d.). Mapping and selection of downy mildew resistance in spinach cv. Whale by low coverage whole genome sequencing. PubMed Central (PMC). https://www.ncbi.nlm.nih.gov/pmc

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